Southeastern Alaska Distribution: New to the district. From a ridge on the west shore of Muir Inlset, roughly opposite Nunatak Knob, alder successional stage, Worley 10552. Annette Island, Hermann 22241.
Habitat: The southeastern Alaska plants grew in a marble cliff crevice. The species is otherwise known from both calcareous rocks and soils from sea level (in the North) to the alpine (in the South).
Comments: Sometimes with sporophytes (the southeastern Alaska plants were sterile). Persson and Weber (1958) note that the species "...seems to avoid more oceanic regions and is perhaps best treated as subcontinental." In western North America the only coastal locality known for the species is the Glacier Bay site.