Lycopodiaceae

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Family: Lycopodiaceae

Group: Vascular Plant, Species

Scientific NameCommon NameSummary
Diphasiastrum alpinumAlpine ClubmossSmall, trailing clubmoss with flattened, 4-ranked leaves infrequently found in high elevation forest and tundra and occasionally at lower elevations. In the region, it has been collected more often in the mountains from the lower part of Lynn Canal north.
Diphasiastrum complanatumGround CedarApparently few collections in the region, the distribution of this species seems to favor areas more influenced by continental climates.
Diphasiastrum sitchenseAlaskan ClubmossTrailing club moss with rounded stems found in high elevation forest and tundra. Collections are primarily from the northern half of the region on mountains near the Gulf of Alaska, though it is unclear whether this represents the actual distribution accurately.
Huperzia continentalisAlpine Fir-mossFairly common larger clubmoss of (sub)alpine tundra where it is the only Huperzia typically found. Formerly considered a part of H. selago. A few scattered collections from across much of the region.
Huperzia miyoshianaPacific Fir-mossA few collections in ARCTOS scattered across the region.
Huperzia occidentalisWestern Fir-mossA single collection in ARCTOS from Port Alexander.
Lycopodiella inundataBog ClubmossKnown from south of Fredrick Sound (including the southern half of Baranof Island), though FNA Range map implies it is to be expeted throughout much of the region. In 2009 a collection was made on Chichagof Island. Can be abundant where found, but reports have been infrequent, with only a few collections from the region.
Lycopodium clavatumRunning ClubmossFound mostly in open areas from low elevation to subalpine. This species may reach the northern extent of its regular range in the region (FNA map), though it is probably fairly common throughout. Collections have been made throughout the region (though see L. lagopus).
Lycopodium dendroideumTree ClubmossExpected throughout the region (FNA range map), collections of this species are mostly from about Frederick Sound south (including mid-Baranof Island).
Lycopodium lagopusOne-cone ClubmossExpected from the far northern mainland part of the region (FNA range map). As of late 2013, only one collection currently in Arctos (from the Juneau Ice Field) - perhaps some older ones are still labeled L. clavatus?
Spinulum annotinumStiff ClubmossCommon throughout the region with many collections. Found in forests and muskegs from sea level to subalpine.

Curtis R. Björk (2020): Notes on the Holarctic Species of Huperzia (Lycopodiaceae), with Emphasis on British Columbia, Canada. _Annales Botanici Fennici_ 57(4-6): 255-278; (28 October 2020). https://doi.org/10.5735/085.057.0410

Abstract Taxonomy of the genus Huperzia (Lycopodiaceae) in western North America and Europe was studied using morphological characters. Delimitations of species were redefined after examining the morphology of the Huperzia selago lectotype and other type specimens in the genus. Two new species were named: H. acicularis Björk from North America and Eurasia, and one species apparently endemic to Europe, H. europaea Björk. Huperzia occidentalis and H. porophylla were found to be conspecific with the Macaronesian endemic H. suberecta (Löwe) Tardieu, which is the oldest name of the three. Additionally, Huperzia suberecta was found to be common and widespread in the northern hemisphere; it, and H. europaea are frequently confused with H. selago.

Key to Huperzia of British Columbia

1. Leaves spreading to reflexed, at least some widest distally ............................................................ H. suberecta

1. Leaves erect to spreading, not reflexed unless damaged, widest proximally or some straight-sided .................... 2

2. Gemmae and gemmaphores scattered and solitary, or if some pseudowhorls present, then numerous gemmae solitary between them; stem foliar outline lacking periodic protrusions, not appearing tiered .......................... 3

2. Gemmae and gemmaphores all or almost all positioned in distinct, evenly spaced (sometimes densely spaced) pseudowhorls that give the stem foliar outline a more or less tiered appearance (N.B.: pseudowhorls may be so close together as to be difficult to discern individually).. ..................................................................................... 4

3. Juvenile stems nodding; leaves more or less subulate; gemmaphores conspicuous, on relatively long stalks ..... ................................................................. H. miyoshiana

3. Juvenile stems erect; leaves narrowly deltate to lanceolate; gemmaphores on short, inconspicuous stalks ......... ............................................................... H. continentalis

4. Gemmaphores protruding well past leaves, pseudowhorls usually closely spaced, and consequently, the stem foliar outline appearing ragged; leaves claw-shaped; plants yellow-green, yellow or orangish ........ H. arctica

4. Gemmaphores not or scarcely exserted, pseudowhorls moderately to widely spaced, and consequently the stem foliar outline appearing distinctly tiered; leaves not distinctly claw-shaped; plants yellow-green or deep green . ...................................................................................... 5

5. Plants yellow-green; leaves acicular, more or less straight and stiffly ascending; gemma pseudowhorls widely distant; gemmae strongly protruding .............. H. acicularis

5. Plants green; leaves narrowly oblong-lanceolate, lanceolate or deltate-lanceolate, variably straight to moderately incurved, ascending to appressed; gemma pseudowhorls moderately distant; gemmae included to weakly protruding ................................................................... H. selago