"?" is not in the list of possible values (Documented, Reported, Unsubstantiated, Expected, Not known to occur, Insufficient Data) for this property.
Southeastern Alaska Distribution: See Map 174. Probably throughout.
Habitat: A pioneer on mineral and semi-coarse substrates, in exposed or shaded sites, usually in wet areas or seeps, but sometimes on well drained materials. Usually in places that are at least somewhat basic. Also on rocks and the bases of trees and shrubs.